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Sokolovskaya, A. P., N. S. Probatova & E. G. Rudyka. 1985. Chromosome numbers in the species of the families Asteraceae, Poaceae, Rosaceae from the Primorye Region, Kamchatka and Sakhalin. Bot. Zhurn. SSSR 70(1): 126–128.
Volkova, S. A. & E. V. Boyko. 1989. Chromosome numbers in representatives of some families of the flora of the Soviet far east. Bot. Žhurn. (Moscow & Leningrad) 74: 1810–1811.
Rudyka, E. G. 1995. Chromosome numbers in vascular plants from the southern part of the Russian Far East. Bot. Žhurn. (Moscow & Leningrad) 80(2): 87–90.
Ito, M., A. Soejima & K. Watanabe. 1998. Phylogenetic relationships of Japanese aster (Asteraceae, Astereae) sensu lato based on chloroplast-DNA restriction site mutations. J. Pl. Res. 111: 217–223.